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Hallmarks of Aging

Primary	Antagonistic	Integrative
Genomic Instability	Cellular Senescence	Stem Cell Exhaustion
Susana Gonzalo, PhD Saint Louis University	James Kirkland, MD, PhD Mayo Clinic	Isabel Beerman, PhD NIA
Vera Gorbunova, PhD University of Rochester	Lu Wang, PhD UT Health San Antonio	Jenny Hsieh, PhD The University of Texas at San Antonio
Telomere Attrition	Mitochondrial Dysfunction	Altered Intercellular Communication
Ronald A. DePinho, MD The University of Texas MD Anderson Cancer Center	Gerald Shadel, PhD Salk Institute	Irina Conboy, PhD UC Berkeley
Epigenetic Alterations	David Marcinek, PhD University of Washington	Chronic Inflammation
Weiwei Dang, PhD Baylor College of Medicine, Houston	Deregulated Nutrient-Sensing	Maxim Artyomov, PhD Washington University in St. Louis
Bess Frost, PhD UT Health San Antonio	Joseph Takahashi, PhD UT Southwestern	Ann Griffith, PhD UT Health San Antonio
Loss of Proteostasis	Dudley Lamming, PhD University of Wisconsin	Dysbiosis
William E. Balch, PhD Scripps Research		Shuo Han, PhD Duke University School of Medicine
Disabled Macroautophagy		Mansour Zadeh, PhD UT Health San Antonio
Rajat Singh, MD Albert Einstein College of Medicine